A couple things about the 305, not sure if the 405 does too: you can change the screens depending on what you want to see when you run. I personally like to see my total time, total distance, and current pace/mile. I have a split screen that shows all 3 at once. You can set heart rate zones and pace zones. You can set timers on it if you want to run intervals.
I upload my runs and then you can create maps that show cross streets. You can create a 'course' whihc is a route you often run, and then you run against a 'virtual partner', that keeps you on pace. It is real motivating to look at your watch and see that your partner is beating you by 30 feet, so you pick up the pace and pull ahead before the finish. After a run I can look at how i did comparing my pace versus my heartrate. It records the elevation, how far you went uphill or downhill.
One great thing about GPS watches is that you can run wherever you want to go and it records your distance, so you don't have to do a set route if you want to avoid traffic or the sun in your eyes or whatever. I am happy with my 305, but it does sometimes take a while to establish the link with the satelites (2-3 minutes on a bad day, but i am usually warming up and stretching a bit anyway).
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